Understanding the Differences Between .223 and 5.56mm Cartridges

Understanding the Differences Between .223 and 5.56mm Cartridges
By WG, July 23, 2025For firearm enthusiasts, hunters, and competitive shooters, the distinction between .223 Remington and 5.56mm NATO cartridges is a topic of frequent discussion. While these two rounds appear nearly identical and are often used interchangeably in rifles like the AR-15, subtle differences in their design, performance, and compatibility can have significant implications. This article breaks down the key differences between .223 and 5.56mm cartridges to help shooters make informed decisions.
Origins and Specifications
The .223 Remington cartridge, introduced in 1962 by Remington Arms, was designed as a commercial round for varmint hunting and target shooting. It became popular due to its high velocity, flat trajectory, and low recoil, making it ideal for small game and precision shooting. The cartridge typically fires a bullet weighing between 40 and 77 grains at velocities ranging from 2,700 to 3,200 feet per second (fps), depending on barrel length and load.
The 5.56x45mm NATO cartridge, developed shortly after in the early 1960s, was created for military use, specifically for the M16 rifle adopted by the U.S. military. While based on the .223 Remington, the 5.56mm NATO round was engineered to meet NATO standards, prioritizing reliability and performance in combat scenarios. It commonly uses a 55- or 62-grain bullet, with velocities around 2,900–3,100 fps from a 20-inch barrel.
Physical and Performance Differences
At first glance, the .223 Remington and 5.56mm NATO cartridges look nearly identical, with the same case dimensions (2.26 inches long) and bullet diameter (.224 inches). However, the differences lie in their internal specifications and performance:
  1. Pressure Levels: The 5.56mm NATO round is loaded to higher pressures than the .223 Remington. The 5.56mm NATO generates chamber pressures around 62,000 PSI (per NATO EPVAT standards), while the .223 Remington is rated at approximately 55,000 PSI (per SAAMI standards). This higher pressure in the 5.56mm results in greater muzzle velocity and energy, but it also places more stress on the firearm.
  2. Chamber and Throat Design: The 5.56mm NATO has a longer throat (or leade) in the chamber, allowing for higher-pressure loads and longer, heavier bullets. The .223 Remington chamber has a shorter throat, optimized for lighter bullets and lower pressures. This difference in chamber design is critical for firearm compatibility.
  3. Bullet Types: While both cartridges can use similar bullet weights, 5.56mm NATO rounds are often loaded with military-style projectiles, such as the M193 (55-grain FMJ) or M855 (62-grain with a steel penetrator core). The .223 Remington is more commonly associated with commercial loads, including varmint-specific rounds with polymer tips or lighter bullets for high velocity.
Firearm Compatibility
One of the most critical considerations for shooters is whether their firearm is chambered for .223 Remington, 5.56mm NATO, or both. Here’s a breakdown of compatibility:
  • Firearms Chambered for .223 Remington: Rifles with a .223 Remington chamber (e.g., some bolt-action rifles or older AR-15s) are designed for the lower-pressure .223 loads. Firing 5.56mm NATO ammunition in a .223 chamber can lead to excessive pressure, potentially causing malfunctions, accelerated wear, or, in rare cases, catastrophic failure. Caution: Always check the barrel markings or consult the manufacturer before using 5.56mm ammo in a .223-chambered firearm.
  • Firearms Chambered for 5.56mm NATO: Rifles designed for 5.56mm NATO, such as most modern AR-15s, can safely fire both 5.56mm and .223 Remington ammunition. The looser 5.56mm chamber accommodates the higher-pressure military round while still functioning reliably with the lower-pressure .223.
  • .223 Wylde Chamber: A popular hybrid chamber, the .223 Wylde is designed to safely fire both .223 Remington and 5.56mm NATO rounds while maintaining excellent accuracy. This chamber splits the difference between the two, offering tighter tolerances than a 5.56mm chamber but more flexibility than a .223 chamber.
Practical Implications for ShootersThe choice between .223 Remington and 5.56mm NATO depends on the shooter’s needs and firearm:
  • Accuracy: The .223 Remington is often favored by precision shooters and varmint hunters due to its lower pressure and availability of specialized loads. Rifles chambered in .223 or .223 Wylde may offer slightly better accuracy with commercial .223 ammo compared to 5.56mm military loads.
  • Availability and Cost: Both cartridges are widely available, but .223 Remington tends to have more variety in commercial loads, while 5.56mm NATO is often cheaper in bulk due to its military surplus availability. Prices fluctuate based on market demand, but as of 2025, .223 Remington rounds typically cost $0.40–$1.00 per round, while 5.56mm NATO can be found for $0.30–$0.80 per round.
  • Intended Use: For self-defense or tactical applications, 5.56mm NATO’s higher velocity and penetration make it a better choice, especially with rounds like the M855 for barrier penetration. For hunting small game or target shooting, .223 Remington’s specialized loads may be more suitable.
Safety Considerations
To avoid potential issues, always verify your firearm’s chambering before selecting ammunition. Barrels are typically marked with the caliber (.223 Rem, 5.56 NATO, or .223 Wylde). If in doubt, consult the manufacturer or a qualified gunsmith. Additionally, inspect ammunition for proper headstamps (.223 REM or 5.56x45mm) to ensure compatibility.
Conclusion
While the .223 Remington and 5.56mm NATO cartridges share many similarities, their differences in pressure, chamber design, and intended use make it essential for shooters to understand their firearm’s capabilities. By choosing the appropriate cartridge for your rifle and purpose—whether it’s precision shooting, hunting, or tactical applications—you can ensure safety, reliability, and optimal performance. Always prioritize safety by confirming your firearm’s chambering and selecting ammunition accordingly.For more information on ammunition and firearm compatibility, consult your firearm’s manual or visit reputable sources like the Sporting Arms and Ammunition Manufacturers’ Institute (SAAMI) or your local gunsmith.
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